Llanelli forward Chris Wyatt has failed to make the Welsh World Cup squad announced today by coach Steve Hansen.
Cardiff pair Jamie Robinson and Ben Evans have also been omitted.
But there are call-ups for Shane Williams, who can play wing or scrum-half; Llanelli wing Garan Evans; and Celtic Warriors lock Brent Cockbain, brother of Australia international Matt.
Hansen has named 31 players and will reduce his current four-strong contingent of hookers to three by the World Cup deadline date of September 10th.
"Chris Wyatt has been very, very unlucky to miss out on a second World Cup," said Hansen. "Naturally, he is disappointed, and we have had a long talk, and I still see him having a future for Wales. Unfortunately, we could only take four locks with us this time.
"The combination we have selected we feel is better suited for this particular tournament.
"Overall, the selection process was a difficult one to go through as, for once, we have more than 30 quality players to choose from.
Hansen said that he would be "addressing the captaincy issue" at a later date.
Wales are in the same qualifying group as New Zealand, Italy, Canada and Tonga Down Under, and are likely to meet England in the quarter-finals if they get that far.
Full squad:
Backs: Kevin Morgan (Celtic Warriors), Rhys Williams (Cardiff Blues), Garan Evans (Llanelli Scarlets), Gareth Thomas (Celtic Warriors), Mark Jones (Llanelli Scarlets), Mark Taylor (Llanelli Scarlets), Tom Shanklin (Cardiff Blues), Sonny Parker (Celtic Warriors), Iestyn Harris (Cardiff Blues), Ceri Sweeney (Celtic Warriors), Stephen Jones (Llanelli Scarlets), Dwayne Peel (Llanelli Scarlets), Gareth Cooper (Celtic Warriors), Shane Williams (Neath-Swansea Ospreys).
Forwards: Jonathan Thomas (Neath-Swansea Ospreys), Martyn Williams (Cardiff Blues), Colin Charvis (unattached), Dafydd Jones (Llanelli Scarlets), Alix Popham (Leeds Tykes), Michael Owen (Gwent Dragons), Gareth Llewellyn (Neath-Swansea Ospreys), Robert Sidoli (Celtic Warriors), Brent Cockbain (Celtic Warriors), Gethin Jenkins (Celtic Warriors), Adam Jones (Neath-Swansea Ospreys), Duncan Jones (Neath-Swansea Ospreys), Iestyn Thomas (Llanelli Scarlets), Gareth Williams (Cardiff Blues), Robin McBryde (Llanelli Scarlets), Huw Bennett (Neath-Swansea Ospreys), Mefin Davies (Celtic Warriors).